台灣菫菜屬物種註記

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菫菜科 Violaceae

目錄

菫菜屬 Viola

最近檢視中國植物誌Viola屬物種,整理中國菫菜屬分類綱要,發現訂正很多物種,陸續下載文獻補齊,台灣植物誌列舉的名錄匯整應紹舜老師的台灣高等植物採色圖誌與臺灣大學植物標本館 植物名彙與基本資訊列舉的同物異名學名,透過The Plant List重新比對學名採用過程,中国植物志哈佛版可以見到部分物種手繪圖,分類模式採用中國植物誌的中文論述,暫時列舉如下,待日後確認。

二裂花柱亞屬 Viola subg. Dischidium

短距堇菜組 Viola Sect. Brevicalcaratae

雙黃花菫菜 Viola biflora

Viola tayemonii Hayata → Viola biflora L.
Viola kanoi Sasaki → Viola biflora L.

美麗堇菜亞屬 Viola subg. Melanium

堇菜亞屬 Viola subg. Viola

合生托葉組 Viola Sect. Adnatae

無地上莖,也無匍匐枝。托葉1/2-2/3與葉柄合生。花柱基部微膝曲,柱頭頂部微凹,兩側具緣邊,前方具短喙。

喜岩菫菜 Viola adenothrix

喜岩菫菜 Viola adenothrix Hayata → 柔毛菫菜 Viola fargesii H. Boissieu

Viola brachycentra Hayata → Viola fargesii H. Boissieu
Viola thrichopoda Hayata
Viola trichopoda Hayata → Viola fargesii H. Boissieu
Viola taiwanensis W. Becker → 台灣菫菜 Viola formosana Hayata
Viola kasanensis auct. non Hayata
Viola royleana auct. non Wallich
Viola rupicola auct. non Elmer → unresolved
Viola effusa W. Becker → unresolved
Viola matsudai Hayata
Viola matsudae Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y.S. Chen & Q.E.Yang

雪山菫菜 Viola adenothrix var. tsugitakaensis

雪山菫菜 Viola adenothrix Hayata var. tsugitakaensis (Masam.) Wang & Huang  →  柔毛菫菜 Viola fargesii H. Boissieu

Viola tsugitakaensis Masam. → 柔毛菫菜 Viola fargesii H. Boissieu

箭葉菫菜 Viola betonicifolia

箭葉菫菜 Viola betonicifolia Sm.

Viola caespitosa D. Don → Viola betonicifolia Sm.
Viola oblongosagittata Nakai var. violascens Nakai → Viola betonicifolia Sm.
Viola oblongosagittata Nakai → 長萼菫菜 Viola inconspicua Blume
Viola patrinii auct. non DC.
Viola patrinii DC. var. chinesis auct. non Ging

短毛菫菜 Viola confusa

短毛菫菜 Viola confusa Champ. ex Benth.  →  長萼菫菜 Viola inconspicua Blume

Viola japonica auct. non Langsdorf
Viola stenocentra Hayata ex Nakai → 琉球菫菜 Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
Viola pseudojaponica Nakai → 琉球菫菜 Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
Viola yedoensis Makino var. pseudojaponica (Nakai) Hashim. ex E. Hama → 琉球菫菜 Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
Viola philippica auct. non Cav.

小菫菜 Viola inconspicua subsp. nagasakiensis

小菫菜 Viola inconspicua Blume subsp. nagasakiensis (W. Becker) Wang & Huang → ???

Viola confusa subsp. nagasakiensis (W. Becker) F. Maek. & Hashim. → 長萼菫菜 Viola inconspicua Blume
Viola mandshurica subsp. nagasakiensis W. Becker → 長萼菫菜 Viola inconspicua Blume
Viola minor (Makino) Makino → 長萼菫菜 Viola inconspicua Blume
Viola patrinii DC. var. minor Makino → 長萼菫菜 Viola inconspicua Blume
Viola taiwaniana Nakai → 琉球菫菜 Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen

紫花地丁 Viola mandshurica

Viola oblongo-sagittata Nakai var. violascens Nakai f. ishizakii Yamamoto → ???
Viola oblongosagittata Nakai var. violascens Nakai → Viola betonicifolia Sm.
Viola oblongosagittata Nakai f. ishizakii Yamam. → Viola mandshurica W. Becker
Viola mandshurica W. Becker var. ciliata Nakai → Viola mandshurica W. Becker
Viola mandshurica W. Becker var. glabra Nakai → Viola mandshurica W. Becker

琉球菫菜 Viola philippica var. pseudojaponica

Viola longistipulata Hayata → Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
南投菫菜 Viola nantouensis S. S. Ying → Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
Viola pseudojaponica Nakai
Viola stenocentra Hayata ex Nakai
Viola taiwaniana Nakai
Viola yedoensis var. pseudojaponica (Nakai) Hashim. ex E. Hama

雙緣柱頭堇菜組 Viola Sect. Bilobatae

有或無地上莖或具上升的匍匐枝。托葉大形呈葉狀或小而近全緣,或具細齒裂,離生或僅基部與葉柄合生。花小;下方花瓣的距較短;花柱基部或多或少膝曲,柱頭頂部兩側有肥厚的片狀緣邊,前方具明顯的短喙;柱頭孔較粗。

如意草 Viola arcuata

Viola verecunda auct. non A. Gray
Viola arcuata Bl. var. verecunda (A. Gray) Nakai → Viola arcuata Blume
Viola arcuata Bl. var. verecunda (A. Gray) Nakai f. rasicans (Makino) Nakai → ???
Viola arcuata f. radicans (Makino) Nakai → Viola arcuata Blume
Viola verecunda A. Gray f. hensoaensis Kudo & Sasaki → Viola arcuata Blume
Viola verecunda A. Gray f. radicans Makino → Viola arcuata Blume

蔓莖堇菜組 Viola Sect. Diffusae

無地上莖,有匍匐枝。匍匐枝具較長的節間,頂端簇生蓮座狀葉。托葉較狹,近膜質。花小;下方花瓣的距較短;柱頭頂部二淺裂或平坦,二裂片間的前方具短喙。

茶匙黃 Viola diffusa

Viola diffusa Ging. var. glabella Boiss
Viola diffusa var. glabella H. Boissieu → Viola diffusa Ging.

台灣菫菜 Viola formosana

Viola tozanensis Hayata → Viola formosana Hayata
Viola kawakamii Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola logistipulata Hayata
Viola longistipulata Hayata → Viola philippica Cav. var. pseudojaponica (Nakai) Y. S. Chen
Viola hypoleuca Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola kawakamii Hayata var. stenopetala Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola arisanensis W. Becker → unresolved
Viola takasagoensis Koidz. → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola formosana Hayata var. tozanensis (Hayata) Hsieh
Viola formosana Hayata var. tozanensis (Hayata) C. F. Hsieh → Viola formosana Hayata

川上氏菫菜 Viola formosana var. kawakamii

川上氏菫菜 Viola formosana Hayata var. stenopetala (Hayata) Wang, Huang & Hashim.

Viola formosana var. stenopetala (Hayata) J. C. Wang, T. C. Huang & T. Hashim. → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola kawakamii Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola kawakamii Hayata var. stenopetala Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola hypoleuca Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola matsudai Hayata
Viola matsudae Hayata → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola takasagoensis Koidz. → Viola formosana Hayata var. kawakamii (Hayata) Y. S. Chen & Q. E. Yang
Viola formosana auct. non Hayata:

台北菫菜 Viola nagasawae

Viola nagasawai C. F. Hsieh → Unresolved

Viola acutilabella Hayata → Viola nagasawae Makino & Hayata
Viola nagasawai Makino & Hayata var. acutilabella (Hayata) Nakai → ???
Viola nagasawai C. F. Hsieh var. acutilabella (Hayata) Nakai → Viola nagasawae Makino & Hayata
台灣植物誌採用Viola nagasawai Makino & Hayata var. nagasawaiThe Plant List 接受的學名是Viola nagasawae Makino & Hayata。

普萊氏菫菜 Viola nagasawai var. pricei

Viola pricei W. Becker → Viola nagasawai C. F. Hsieh var. pricei (W. Becker) Wang & Huang → Unresolved
Viola nagasawai auct. non Makino & Hayata

心葉茶匙黃 Viola tenuis

心葉茶匙黃 Viola tenuis Benth. → 茶匙黃 Viola diffusa Ging.

Viola diffusa subsp. tenuis (Benth.) W. Becker → 茶匙黃 Viola diffusa Ging.

斜柱頭堇菜組 Viola Sect. Plagiostigma

無地上莖。根狀莖細長且橫生,或短縮且垂直。托葉離生,通常膜質。柱頭頂部扁平、傾斜,呈釘帽狀,前方具短喙,喙端稍細,有較小的柱頭孔。

尖山菫菜 Viola senzanensis

匍匐堇菜組 Viola Sect. Serpentes

無地上莖,或有時具縮短的地上莖。匍匐莖纖細而伸長,有均勻散生的葉片。花較小;下方花瓣的距較短;柱頭頂部平坦無緣邊或兩側有緣邊,前方具短喙。

新竹菫菜 Viola kosanensis

新竹菫菜 Viola shinchikuensis Yamamoto

Viola shinchikuensis Yamam. → Viola kosanensis Hayata
Viola nagamiana Hashimoto
Viola nagamiana T. Hashim. → Viola kosanensis Hayata

鳥嘴柱頭組 Viola Sect. Trigonocarpae

花期有地上莖,稀無地上莖。柱頭前方的鉤狀喙較短,喙通常向上斜升,稀近直立,較花柱最寬部分的直徑為短,柱頭孔較粗。蒴果長圓形;果梗通常直立。

雞腿菫菜 Viola acuminata

紫花菫菜 Viola grypoceras

Viola acuminata Ledcbour → Viola acuminata Ledeb.

翠峰菫菜 Viola obtusa var. tsuifengensis

Violaceae Reference

  1. Dani, Ram S and Krishna K Shrestha 2004 Two new records of Viola L. (Violaceae) for Nepal Him J Sci 2 (3): 48-50
  2. Freitas leandro and Marlies Sazima 2003 Floral Biology and Pollination Mechanisms in Two Viola Species—From Nectar to Pollen Flowers? Ann Bot 91(3): 311-317
  3. Hashimoto, Tamotsu 1987 Trigonocarpae Violets of Taiwan Annals of the Tsukuba Botanical Garden 6 1-7
  4. Ning, Zu-Lin, Zhen-Xin Zeng, Ling Chen, Bing-Qiang Xu and Jing-Ping Liao 2012 Viola jinggangshanensis (Violaceae), a new species from Jiangxi, China Annales Botanici Fennici 49 (5/6) 383-386
  5. Peng, Ching-I and Yueh-Fong Chen 1985 Hybanthus Jacq. (Violaceae), a New Generic Record for the Flora of Taiwan Bot. Bull. Acad. Sin. 26 (2) 213-220
  6. Huang, Su-Hua 2003 New Species of Viola from Yunnan Acta Botanica Yunnanica 25(4) 431-434
  7. Wang, Jenn-Che, Tseng-Chieng Huang 1990 Notes on the Flora of Taiwan (9) --- The Viola L. (Violaceae) Taiwania 35(1) 12-56
  8. Zhou, Jin-Song and Fu-Wu Xing 2007 Viola changii sp. nov. (Violaceae) from Guangdong, southern China Nordic Journal of Botany 25: 303-305

  1. 王庆瑞 1991 中国堇菜属植物新分类群 云南植物研究 13(3): 257-266
  2. 王庆瑞 1992 中国堇菜属植物分类群(续) 云南植物研究 14(4) 379-382
  3. 周勁松、邢福武 2007 中國堇菜屬美麗堇菜亞屬(堇菜科)植物分類學修訂 西北植物學報 27(5) 1017-1021
  4. 周劲松、龚琴、刑福武 2007 中国堇菜(堇菜科)两新异名 热带亚热带植物学报 15(4):366-368
  5. 周劲松、巫远坤、黎昌汉、邢福武 2007 广东堇菜属植物新记录 热带亚热带植物学报 15(5):455-456
  6. 李连方、尹祖棠 2000 堇菜属植物花粉及叶表皮超微形态多样性的研究 中国农业大学学报 5(3) : 6-12
  7. 邢福武、秦新生 2004 中国植物志堇菜科 Violaceae拾遗 华南农业大学学报(自然科学版) 25(1) 120-121
  8. 邢福武、黎昌汉 2004 堇菜属 (堇菜科)植物新资料 热带亚热带植物学报 12(5):475-476
  9. 邢福武, 秦新生 2004 中國植物志菫菜科Violaceae拾遺 華南農業大學學報 25(1) 120-121
  10. 孙雪芹,刘玫,孙天航,张欣欣,史传奇 2014 东北堇菜属植物叶形态结构的研究及其分类学价值的探讨 草业学报 23 (2): 223-234
  11. 陳又生 2006 中国堇菜属的分类修订 中國科學院研究生院(植物研究所)
  12. 梁冠欣,邢福武 2010 堇菜属组间的系统发育与亲缘关系 : 基于trn L-trn F、psbA-trnH、 rpL16、ITS序列, 细胞学以及形态学证据 云南植物研究 32(6): 477-488
  13. 黄素华 2003 云南堇菜属新种 云南植物研究 25(4): 431-434
  14. 潘雅书,邢福武 2012 长萼堇菜多倍体复合体种群的分布格局研究 植物科学学报 30(2) 147-152
  15. 龚琴、周劲松、张永夏、梁冠欣、陈红锋、邢福武 2010 国产堇菜属的分子系统学研究 热带亚热带植物学报 18(6):633-642
  16. 藤原 正人 and 笠原 恵 スミレ属の葉緑体 DNA を用いた分子系統解析に関する研究
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