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Phyllanthaceae Reference

  1. Airy Shaw, H. K. 1980 New or noteworthy Australian Euphorbiaceae II. Muelleria 4: 207-241.
  2. Barker, Christine and P.C. Van Welzen 2010 Flueggea (Euphorbiaceae s. l. or Phyllanthaceae) in Malesia Systematic Botany 35 (3):541-551
  3. Croizat, Leon and Hiroshi Hara 1940 Critical notes on some Formosan Euphorbiaceae (I) J. Jap. Bot. 16 (6): 313–326
  4. Croizat, Leon and Hiroshi Hara 1940 Critical notes on some Formosan Euphorbiaceae (II) J. Jap. Bot. 16 (7): 383–390
  5. Dressler, Stefan 1996 The genus Bridelia (Euphorbiaceae) in Malesia and Indochina - A regional revision Blumea 41: 263-331
  6. Heijkoop1, M. and P. C. van Welzen 2017 A revision of the genus Actephila (Phyllanthaceae) in the Malesian region Blumea 62 7–25
  7. Hoffmann, Petra 2000 Checklist of the genus Antidesma (Euphorbiaceae) in Thailand Thai. for. Bull. (Bot). 28 139-156
  8. Hurusawa, Isao 1947 Spicilegium plantarum Asiae Orientalis I Bot. Mag. (Tokyo) 60 (703-714): 71-76
  9. Jung, Ming-Jer and Ching-Long Yeh 2010 Some supplements on the Antidesma pleuricum Tul. (Euphorbiaceae), a neglected species in the Flora of Taiwan Taiwania 55(3): 318-323
  10. Keng, Hsuan 1955 The Euphorbiaceae of Taiwan Taiwania 6 (1): 27-66
  11. Kurosawa, Takahide 2006 The Family Euphorbiaceae s. l. in Nepal A Checklist and Taxonomical Notes Univ. Mus. Univ. Tokyo Bull. 42: 155-346
  12. Kurosawa, Takahide 2016 Transfer of Japanese Breynia and Glochidion to Phyllanthus (Phyllanthaceae) J. Jpn. Bot. 91: 118–121
  13. Low, Y. W., P. K. F. Leong, S. P. Tee, Rajesh Singh, M. L. C. Tay and K. M. Wong 2014 Margaritaria (Phyllanthaceae), a new generic record for the Singapore flora Gardens' Bulletin Singapore 66(1) 47–56
  14. Schot, Anne M. 1995 A synopsis of taxonomic changes in Aporosa Blume (Euphorbiaceae) Blumea 40 449-460
  15. Schot, Anne M. 1998 Biogeography of Aporosa (Euphorbiaceae): testing a phylogenetic hypothesis using geology and distribution patterns Biogeography and Geological Evolution of Se Asia 279-290
  16. Schot, Anne Marketta 2004 Systematics of Aporosa (Euphorbiaceae) Blumea Supplement 17
  17. Van Welzen, Peter C. 2003 Revision of the Malesian and Thai species of Sauropus (Euphorbiaceae: Phyllanthoideae) Blumea 48(2) 319-391
  18. Van Welzen, Peter C. and Kanchana Pruesapan 2010 Four new species of Breynia (Phyllanthaceae / Euphorbiaceae sensu lato) and one new combination from Thailand and Malaysia THAI FOR. BULL. (BOT.) 38: 111–119
  19. Van Welzen, P. C., K. Pruesapan, I. R. H. Telford, H.-J. Esser, J. J. Bruh 2014 Phylogenetic reconstruction prompts taxonomic changes in Sauropus, Synostemon and Breynia (Phyllanthaceae tribe Phyllantheae) Blumea 59: 77–94
  20. Wurdack, Kenneth J., Petra Hoffmann, Rosabelle Samuel, Anette de Bruijn, Michelle van der Bank, Mark W. Chase. 2004 Molecular phylogenetic analysis of Phyllanthaceae (Phyllanthoideae pro parte, Euphorbiaceae sensu lato) using plastid RBCL DNA sequences Am J Bot. 91(11): 1882-1900
  21. Zhang, Jing, Shuxia Wang, Houhun Li, Bingbing Hu, Xiaofei Yang, Zhibo Wang 2012 Diffuse Coevolution between Two Epicephala Species (Gracillariidae) and Two Breynia Species (Phyllanthaceae) PLoS ONE 7 (7): e41657

  1. 李永泉,张奠湘 2007 山地五月茶的蝇类传粉研究 植物分类学报 45(2): 217–226
  2. 李永泉,罗中莱,张奠湘 2007 叶下珠科花粉组织化学、花粉数和胚珠数及其与传粉者关系的研究 生物多样性 15(6): 645–651
  3. 李秉滔 1988 中国叶下珠亚科植物资料 植物分类学报 26 (1): 58-65

Glochidion Reference

  1. Chakrabarty, T. & Gangopadhyay, M. 1995 The genus Glochidion (Euphorbiaceae) in Indian subcontinent J. Econ. Taxon. Bot. 19: 173-234.
  2. Chia, Edmund J. J., Z. Y. Ooi, H. K. Lua, Adrian H. B. Loo, W. F. Ang, K. H. Ong and K. M. Wong 2017 Glochidion obscurum Bl. (Phyllanthaceae), a new record for Singapore NATURE IN SINGAPORE 10: 49–53
  3. Croizat, Leon and Hiroshi Hara 1940 Critical notes on some Formosan Euphorbiaceae (I) J. Jap. Bot. 16 (6): 313–326
  4. Hatusima, Sumihiko 1956 New and Noteworthy Plants from the Ryukyu Islands and Formosa Science bulletin of Agriculture & Home Economics Division, University of the Ryukyus 3: 19 -33
  5. Kato, Makoto, Atsushi Takimura and Atsushi Kawakita 2003 An obligate pollination mutualism and reciprocal diversification in the tree genus Glochidion (Euphorbiaceae) PNAS 100 (9): 5264-5267
  6. Kurosawa, Takahide 2016 Transfer of Japanese Breynia and Glochidion to Phyllanthus (Phyllanthaceae) J. Jpn. Bot. 91: 118–121
  7. Lee, Chung Suk and Hyoung Tak Im 1994 Glochidion chodoense C. Lee et Im (Euphobiaceae) , a new species from Chodo Archipelago, Korea Korean J. Pl. Taxon. 24 (1):13-16
  8. Rao, M. Sankara, J. Swamy. S. Nagaraju,S. B. Padal,M. Tarakeswara Naidu, K. Chandramohan, T. Thulasiah 2016 Glochidion talakonense sp. nov. (Phyllanthaceae) from Seshachalam Biosphere Reserve, Andhra Pradesh, India Bangladesh J. Plant Taxon. 23 (1): 59-63
  9. Rasingam, L., Alok R. Chorghe, P.V. Prasanna and M. Sankara Rao 2014 Glochidion tirupathiense (Phyllanthaceae) ─ A New Species from Seshachalam Biosphere Reserve of Andhra Pradesh, India Taiwania 59 (1): 9–12
  10. Sandhya, S., Chaintanya R. S. N. A. K. K, Vinod K. R., K. N. V. Rao, David Banji, Sudhakar K and Swetha R 2010 An updated review on the Genus Glochidion Plant Archives of Applied Science Research 2 (2):309-322
  11. Welzen, Peter C. van, R. M. A. P. Haegens, J. W. F. Slik, S. M. Bollendroff, S. Dressler and H.-J. Esser 2000 Checklist of the genera of Thai Euphorbiaceae - I Thai For. Bull. (Bot.) 28: 59-111
  12. Yao, Gang and Dian-Xiang Zhang 2013 Reinstatement of three names in Glochidion (Phyllanthaceae) based on morphological and palynological evidence Nordic Journal of Botany 31: 001–007
  13. Yao, Gang and Dianxiang Zhang 2015 Taxonomic notes on Glochidion acuminatum and G. triandrum (Phyllanthaceae) Phytotaxa 236 (1):79-85
  14. Yao, Gang and Dian‐Xiang Zhang 2016 Pollen morphology of Chinese Glochidion (Phyllanthaceae) and its taxonomic implications Nordic Journal of Botany 34: 102–110
  15. Yao, Gang, Zhuqiu Song, Youheng Wu, Nguyen Thi Kim Thanh, Yuling Li, Shi Shi and Shixiao Luo 2018 Taxonomic studies of Glochidion (Phyllanthaceae) from the Indo-China Peninsula (II): The identities of G. anamiticum and G. annamense PhytoKeys 114: 1–9
  16. Yao, Gang, Zhu-Qiu Song, Bin-E Xue, Shi Shi, Yu-Ling Li, Shi-Xiao Luo 2020 Taxonomic revision of the genus Glochidion (Phyllanthaceae) in Taiwan, China PhytoKeys 159: 137–159
  17. Zhang, Dianxiang & & Gang Yao 2015 Reinstatement of three names in Glochidion (Phyllanthaceae) based on morphological and palynological evidences Nordic Journal of Botany 33(2):197-203

  1. 呂福原、許媖素 2003 臺灣大戟科之一新種-卵葉饅頭果 林業研究季刊 25(4): 83-86
  2. 許媖素、呂福原、鄧書麟、廖政麒 2006 台灣饅頭果屬植物 (大戟科) 之分類研究 林業研究季刊 28(3) 49-68
  3. 姚纲,张连婕,薛彬娥,罗世孝 2017 中国算盘子属(叶下珠科)果实形态特征及其分类学意义 植物科学学报 35(2): 139-151
  4. 姚纲,张连婕,薛彬娥 2017 中国算盘子属(叶下珠科)一些种的分类学处理 广西植物 37 (5): 547-553

Phyllanthus Reference

  1. Bouman, Roderick W., Paul J. A. Keβler, I. R. H. Telford, J. J. Bruhl, Peter C. van Welzen 2018 Subgeneric delimitation of the plant genus Phyllanthus (Phyllanthaceae) Blumea 63 : 167–198
  2. Bouman, Roderick W., Paul J. A. Keβler and Peter C. van Welzen 2019 Lectotypification and amended description of Phyllanthus (Phyllanthaceae) species described by Koorders from Sulawesi, Indonesia Reinwardtia 18 (2): 97−103
  3. Chantaranothai, Pranom 2005 Taxonomic Notes on the genus Phyllanthus L. (Euphorbiaceae) in Thailand Thai Forest Bull. (Bot.) 33: 16-20
  4. Chen, Shih-Huei and Ming-Jou Wu 1997 A revision of the Herbaceous Phyllanthus L. (Euphorbiaceae) in Taiwan. Taiwania 42(3) 239-261
  5. Chen, Y.-J., S.-H. Chen , T.-C. Huang and M.-J. Wu 2009 Pollen morphology of Philippine species of Phyllanthus (Phyllanthaceae, Euphorbiaceae s. l.) Blumea 54: 47-58
  6. da Silva, Marcos José and Margareth Ferreira de Sales 2008 Sinopse do gênero Phyllanthus (Phyllanthaceae) no Nordeste do Brasil Rodriguésia 59(2) 477-422
  7. Croizat, Leon 1940 Notes on New and Critical Far Eastern Phyllanthus J.. Jap. Bot. I6: 646-658
  8. Dnyaneshwar, Warude, Chavan Preeti, Joshi Kakpana and Patwardhan Bhushan 2006 Development and Application of RAPD-SCAR Marker for Identification of Phyllanthus emblica L. Biol. Pharm. Bull. 29 (11): 2313-2316
  9. Esser, Hans-Joachim 2002 A revision of Triadica Lour. (Euphorbiaceae) Harvard Papers in Botany 7(1): 17-21
  10. Ganesan, R. 2003 Identification, distribution and conservation of Phyllanthus indofischeri, another source of Indian gooseberry CURRENT SCIENCE 84 (12): 1515-1518
  11. Jayasinha, Purniam 1999 Pitawakka-Phyllanthus debilis Medicinal and Aromatic Plant Series No. 10 ISBN 955-8394-07-06
  12. Kurosawa, Takahide 2001 Taxonomy and Distribution of Japanese Phyllanthus (Euphorbiaceae) Acta Phytotax. Geobot. 52 (1): 11-33
  13. Lee, Chao-ying, Shih-wei Tsai, Shau-ting Chiu, Chao-lin Kuo, Chung-chuan Chen and Tai-hui Chiu 2005 The Anatomical Studies of Phyllanthus Species in Taiwan J. Chin. Med. 16 (2-3) 119-148
  14. Luo Shi-Xiao, Hans-Joachim Esser, D. Zhang and S. S. Renner 2011 Nuclear ITS sequences help disentangle Phyllanthus reticulatus (Phyllanthaceae), an Asian species not occurring in Africa, but introduced to Jamaica Syst. Bot. 36: 99–104
  15. Raju, Vatsavaya S., S. Suthari, A. Ragan 2011 The identity and occurrence of Phyllanthus hookeri Muell.-Arg. and P. nozeranii Rossignol & Haicour (Euphorbiaceae) in India Bangladesh J. Plant Taxon. 18 (1): 57-63
  16. Rossignol, L., M. Rossignol and R. Haicour 1987 A Systematic Revision of Phyllanthus Subsection Urinaria (Euphorbiaceae) American Journal of Botany 74(12) 1853-1862
  17. Sangeetha, N., S. Mercy, M. Kavitha, Divya Selvaraj, R. Sathishkumar and D. Ganesh 2010 Morphological variation in the Indian gooseberries ( Phyllanthus emblica and Phyllanthus indofischeri) and the chloroplast trnL (UAA) intron as candidate gene for their identification Plant Genetic Resources 8 (2): 1-7
  18. Silva, Marcos José da and Margareth Ferreira de Sales 2007 Phyllanthus L. (Phyllanthaceae) em Pernambucao, Brasil Acta Bot. Bras. 21 (1): 79-98.
  19. Theerakulpisut, Piyada, Nanawapee Kanawapee, Duangkamol Maensiri, Sumontip Bunnag and Pranom Chantaranothai 2008 Development of species-specific SCAR markers for identification of three medicinal species of Phyllanthus Journal of Systematics and Evolution 46(4): 614–621
  20. Verwijs, I. I. M., Roderick W. Bouman and Peter C. van Welzen 2019 A taxonomic revision of Phyllanthus subgenus Macraea (Phyllanthaceae) Blumea 64: 231-252
  21. Webster, Grady L. 1970 A revision of Phyllanthus (Euphorbiaceae) in the continental United States Brittonia 22 44-76
  22. Webster, Grady L. 1986 A Revision of Phyllanthus (Euphorbiaceae) in Eastern Melanesia Pacific Science 40 1-4
  23. Wu, Ming-Jou, Tseng-Chieng Huang, Chia-Chin Liu, Yeng-Juei Chen, Yung-Sheng Chang, Chia-Lun Hsu, Shin-Yi Wu, Ai-Yi Tseng, Yin-Chen Chang, Cheng-Chuan Liu and Anusara Kaewmuana 2016 Pollen Morphology and Taxonomy in Malesian Phyllanthus (Phyllanthaceae) J. Jpn. Bot. 91 Suppl.: 257–292

  1. 金存礼 1981 中国叶下珠属新植物及一些混淆种的研究 植物分类学报 19 (3): 345-350.
  2. 李秉滔 1987 中国叶下珠属植物的修订 植物分类学报 25 (5): 371-383.
  3. 胡喻华 2006 叶下珠属植物新资料 华南农业大学学报 27(2) 121-124
  4. 韦仲新、丘华兴、周丽华 2002 大戟科叶下珠属花粉形态的研究 云南植物研究 24(2) 253-259
  5. 黒沢高秀 2001 ナガエコミカンソウ(トウダイグサ科)の日本へ帰化 植物研究雑誌 76: 51-52


Web Link

  1. Department of Botany & the U.S. National Herbarium 2017 The Plant Press vol 20 (2) Smithsonian National Museum of Natural History
  2. International Association for Vegetation Science 2009 Vegetation Processes and Human Impact in a Changing World 52nd INTERNATIONAL SYMPOSIUM
  3. 17th International Workshop European Vegetation Survey 2008 Using phytosociological data to address ecological questions Masaryk University, Brno, Czech Republic
  4. Gilly, Charles Louis 1950 Phylogenetic development of the inflorescence and generic relationships in the Kobresiaceae Retrospective Theses and Dissertations 13679.
  5. Acevedo-Rodríguez, P. et al, 2015 Lianas and climbing plants of the Neotropics

Books and Journals


  1. Arora, R. K. 2014 Diversity in underutilized plant species - an Asia-Pacific perspective Bioversity International
  2. Azan, Shakira Stephanie Elaine 2011 Invasive aquatic plants and the aquarium and ornamental pond industries Theses and dissertations Ryerson University P. 818
  3. Beech, E., M. Rivers, S. Oldfield and P. P. Smith 2017 Global Tree Search: The first complete global database of tree species and country distributions Journal of Sustainable Forestry 36: 5 454-489
  4. Bechtel, Albert Reiff 1921 The Floral Anatomy of the Urticales American Journal of Botany 8 (8): 386-410
  5. Chang, C. S., H. Kim, and K. S. Chang. 2014. Provisional Checklist of Vascular Plants For The Korea Peninsula Flora (KPF). Designpost, Pajo.
  6. Chang, C. S., H. Kim and K. S. Chang. 2015. A Catalogue of Vascular Plant Type Specimens from Korea Designpost, Pajo.
  7. Chang, Ching-en 1981 An Eumeration of the Woody Plants of Botel Tobago J. Phytogeogr. Taxon. 29 (1): 1-21
  8. Chen, Xin, Hai He and Li-Bing Zhang 2015 A monograph of the Anisophylleaceae (Cucurbitales) with description of 18 new species of Anisophyllea Phytotaxa 229 (1):1-189
  9. Chiou, Chyi-Rong, Guo-Zhang Michael Song, Jui-Han Chien, Chang-Fu Hsieh, Jenn Che Wang, Ming-Yih Chen, Ho-Yih Liu, Ching-Long Yeh, Yue-Joe Hsia, and Tze-Ying Chen 2010 Altitudinal distribution patterns of plant species in Taiwan are mainly determined by the northeast monsoon rather than the heat retention mechanism of Massenerhebung Botanical Studies 51: 89-97
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  18. Editorial Commitee fo the Flora of Taiwan 1993 Flora of Taiwan, Second edition. Volume Three, Taiwan University, Taipei, Taiwan 106, the Republic of China, ISBN 957-9019-92-4
  19. Editorial Commitee fo the Flora of Taiwan 1998 Flora of Taiwan, Second edition. Volume Four, Taiwan University, Taipei, Taiwan 106, the Republic of China, ISBN 957-9019-92-4
  20. Editorial Commitee fo the Flora of Taiwan 2000 Flora of Taiwan, Second edition. Volume Five, Taiwan University, Taipei, Taiwan 106, the Republic of China, ISBN 957-9019-92-4
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  22. Endress, Peter K; Lorence, David H 2020 Inflorescence structure in Laurales - stable and flexible patterns International Journal of Plant Sciences 181 (3):267-283
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  24. Fosberg, F. Raymond and Sachet, Marie-Hélène 1980 Systematic Studies of Micronesian Plants Smithsonian Contributions to Botany 1–40
  25. Fridley, Jason D. 2008 Of Asian Forests and European Fields: Eastern U.S. Plant Invasions in a Global Floristic Context PLoS ONE 3 (11):e3630
  26. Giesen, Wim Stephan Wulffraat, Max Zieren and Liesbeth Scholten 2006 Mangrove Guidebook for Southeast Asia FAO and Wetlands International ISBN: 974-7946-85-8
  27. Goldberg, Aaron 1986 Classification, Evolution, and Phylogeny of the Families of Dicotyledons Smithsonian Contributions to Botany 58: 1–314
  28. Goldberg, Emma E., Sarah P. Otto, Jana C. Vamosi, Itay Mayrose, Niv Sabath, Ray Ming, and Tia-Lynn Ashman 2017 Macroevolutionary synthesis of flowering plant sexual systems Evolution 71 (4): 898–912
  29. Gradstein, S. Robbert, Jürgen Homeier and Dirk Gansert 2008 The Tropical Mountain Forest Patterns and Processes in a Biodiversity Hotspot Biodiversity and Ecology Series 2
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