台灣懸鉤子屬物種註記

山羊百科，林文智老師的植物、山岳、攝影紀錄

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薔薇科　Rosaceae

懸鉤子屬　Rubus

目錄

矮生莓組　Rubus Sec. Chamaebatus　／單葉

刺萼寒莓　Rubus pectinellus

Rubus pectinellus Maxim. var. trilobus Koidz. → Rubus pectinellus Maxim.

玉山懸鉤子　Rubus pentalobus

玉山懸鉤子　Rubus pentalobus Hayata → Rubus rolfei J. E. Vidal
玉山懸鉤子　Rubus calycinoides Hayata ／ Rubus calycinoides Hayata ex Koidz. → Rubus rolfei J. E. Vidal
大葉玉山懸鉤子　Rubus calycinoides Hayata var. macrophyllus Li → Rubus rolfei J. E. Vidal
Rubus hayata-koidzumii Naruh. → unresolved
Rubus rolfei auct. non S. Vidal
高山懸鉤子　Rubus rolfei Vidal → Rubus rolfei J. E. Vidal

高山懸鉤子　Rubus rolfei

Rubus rolfei Vidal var. lanatus Hayata → Rubus rolfei J. E. Vidal

刺毛莓组 Sect. Dalibardastrum　／單葉

周毛懸鉤子　Rubus amphidasys

空心莓亞屬 Rubus Subgenus Idaeobatus　／ 單、複葉

球果组 Rubus Subgenus Idaeobatus Sect. Corchorifolii

變葉懸鉤子　Rubus corchorifolius

台東刺花懸鉤子　Rubus taitoensis

臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11801
刺花懸鉤子　Rubus aculeatiflorus Hayata → Rubus taitoensis Hayata var. aculeatiflorus (Hayata) H. Ohashi & C. F. Hsieh
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11802
刺花懸鉤子　Rubus mingetsensis Hayata → Rubus taitoensis Hayata var. aculeatiflorus (Hayata) H. Ohashi & C. F. Hsieh
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11801
Rubus aculeatiflorus Hayata var. taitoensis (Hayata) Liu & Yang
Rubus aculeatiflorus var. taitoensis (Hayata) T. S. Liu & T. Y. Yang → Rubus taitoensis Hayata
Rubus taitoensis Hayata var. aculeatiflorus (Hayata) H. Ohashi & Hsieh → Rubus taitoensis Hayata var. aculeatiflorus (Hayata) H. Ohashi & C. F. Hsieh



  • Rubus aculeatiflorus Hayata var. taitoensis (Hayata) Liu & Yang, 12: 12. 1969. (Sci. Ann. Taiwan Prov. Mus., 台灣省立博物館科學年刊)；



  • Rubus taitoensis Hayata var. aculeatiflorus (Hayata) H. Ohashi & Hsieh, Flora of Taiwan, second edition 3: 143. 1993. (Fl. Taiwan 2nd edit., 台灣植物誌第二版)



刺花懸鉤子　Rubus taitoensis var. aculeatiflorus

苦懸鉤子　Rubus trianthus

傘房序組　Rubus Sect. Idaeanthi


白絨懸鉤子　Rubus incanus

白絨懸鉤子　Rubus incanus Sasaki ex T.-S. Liu & T.-Y. Yang → 白絨懸鉤子　Rubus niveus Thunb.
Rubus niveus auct. non Thunb.

柔毛葉組　Rubus Sect. Pungentes

紅花懸鉤子　Rubus inopertus

李棟山懸鉤子　Rubus ritozanensis Sasaki → Rubus inopertus (Focke) Focke
Rubus fraxinifolius Poir. var. yushuni Suzuki & Yamamoto
Rubus fraxinifolius var. yushunii Suzuki & Yamam. → Rubus inopertus (Focke) Focke → unresolved
Rubus yushuni (Suzuki & Yamamoto) Suzuki & Yamamoto ／ Rubus yushuni (Suzuki & Yamam.) Suzuki & Yamam. → Rubus inopertus (Focke) Focke
Rubus ohwianus Koidz. → unresolved ／ In Acta Phytotax. & Geobot. 8: 108. 1939.；https://www.jstage.jst.go.jp/article/bunruichiri/8/2/8_KJ00002594549/_pdf/-char/ja
Rubus yamamotoanus Li → unresolved



  • Rubus ritozanensis Sasaki, Transactions of the Natural History Society of Formosa 21: 250. 1931. (Trans. Nat. Hist. Soc. Formos., 台灣博物學會會報)
  • Rubus ritozanensis Sasaki；劉業經，呂福原，歐辰雄 1994 臺灣樹木誌 169
  • Rubus fraxinifolius Poir. var. yushuni Suzuki & Yamamoto, Transactions of the Natural History Society of Formosa 22: 409. 1932. (Trans. Nat. Hist. Soc. Formos., 台灣博物學會會報)
  • Rubus yushuni (Suzuki & Yamamoto) Suzuki & Yamamoto, Transactions of the Natural History Society of Formosa 25: 130. 1935. (Trans. Nat. Hist. Soc. Formos., 台灣博物學會會報)
  • Rubus ohwianus Koidz., Acta Phytotax. Geobot. 8: 110. 1939.／Koidzumi, Geniti 1939 Contributiones ad Cognitionem Florae Asiae Orientalis (continued from vol. VIII p. 61) Acta Phytotax. Geobot. 8 (2): 108-117


  • Rubus yamamotoanus Li, Woody Flora of Taiwan 319. 1963. (Woody Fl. Taiwan, )


Rubus ritozanensis Sasaki；http://ylist.info/ylist_detail_display.php?pass=41944


毛刺懸鉤子　Rubus pungens var. oldhamii

Rubus oldhamii Miq. → Rubus pungens Cambess.
Rubus pungens auct. non Camb.

刺懸鉤子　Rubus pungens

虎婆刺　Rubus parvipungens Hayata → unresolved
毛刺懸鉤子　Rubus pungens Camb. var. oldhamii (Miq.) Maxim.

台灣莓　Rubus taiwanicolus

Rubus taiwanicolus Koidz. & Ohwi → Rubus taiwanicola Koidz. & Ohwi

Rubus elegans Hayata → Rubus taiwanicola Koidz. & Ohwi
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11803
Rubus takasagomontanus Hatusima ／ Rubus takasagomontanus Hatus. → unresolved

鬼懸鉤子　Rubus wallichianus

Rubus wallichianus Wight & Arnott → Rubus wallichianus Wight & Arn.

Rubus fasciculatus Duthie → Rubus wallichianus Wight & Arn.
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11803
Rubus ellipticus Seem. var. fasciculatus (Duthie) Masam. →
Rubus ellipticus Seem. ssp. fasciculatus (Duthie) Focke → Rubus wallichianus Wight & Arn.
Rubus pinfaensis Lévl. & Van. → Rubus wallichianus Wight & Arn.

多心皮組　Rubus Sect. Rosaefolii

榿葉懸鉤子　Rubus alnifoliolatus

紅狹葉懸鉤子　Rubus cardotii

Rubus fraxinifoliolus Hayata → Rubus fraxinifoliolus Hayata
Rubus rosaefolius Smith var. formosanus Cardot →
Rubus rosaefolius Sm. → Rubus rosifolius Sm.
紅狹葉懸鉤子　Rubus rubro-angustifolius Sasaki → Rubus croceacanthus H. Lév. var. glaber Koidz. 禿懸鉤子
Rubus asper Wall. var. glaber (Koidz.) Hsieh
Rubus asper Wall. ex D. Don → unresolved
Rubus croceacanthus H. Lév. var. glaber auct. non Koidz.

虎婆刺　Rubus croceacanthus

Rubus tagallus auct. non Cham. & Schl.
虎梅刺　Rubus tagallus sensu Matsum. in Bot. Mag. Tokyo 16: 4. 1902, non Cham. & Sohl.
毛刺懸鈎子　Rubus rosaefolius Smith var. hirsutus Hayata → https://tai2.ntu.edu.tw/ebook/ebookpage.php?volume=0&book=Fl.%20Mont.%20Formos.&page=81
毛刺懸鈎子　Rubus hirsutus (Hayata) Hayata → unresolved ／ Rubus hirsutus Thunb. (non)
長果懸鉤子　Rubus asper Wall. ex D. Don var. pekanius Focke → ???
Rubus sorbifolius auct. non Maxim.
圓果懸鉤子　Rubus sphaerocephalus Hayata → Rubus croceacanthus H. Lévl.
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11806
虎梅刺／薄瓣悬钩子　Rubus piptopetalus Hayata ex Koidz. → Rubus croceacanthus H. Lévl.
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11806
長果懸鉤子　Rubus dolichocephalus Hayata → 紅腺懸鉤子　Rubus sumatranus Miq.
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11805
Rubus glanduloso-calycinus Hayata → 腺萼懸鉤子　Rubus glandulosocalycinus Hayata
Rubus parvipungens Hayata → unresolved
毛刺懸鈎子　Rubus hirsutopungens Hayata → unresolved
虎梅刺　Rubus euphlebophyllus Hayata → Rubus croceacanthus H. Lévl.
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11805
Rubus rosaefolius Smith var. polyphyllarius Cardot ／ Rubus rosifolius Sm. var. polyphyllarius Cardot → ???
長果懸鉤子　Rubus somai Hayata → unresolved
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11805
Rubus polyphyllarius (Cardot) Koidz. → unresolved
高砂懸鉤子　Rubus takasagoensis Koidz. → unresolved
Rubus hayatanus Koidz. → 腺萼懸鉤子　Rubus glandulosocalycinus Hayata
Rubus croceacanthus Lévl. var. glaber Koidz. → 禿懸鉤子　Rubus croceacanthus H. Lév. var. glaber Koidz.
Rubus hayatai Nemoto ex Makino ／ 毛刺懸鈎子　Rubus hayatai Nemoto → unresolved
Rubus asper Wall. ex D. Don → unresolved
毛刺懸鈎子　Rubus okamotoanus Koidz. → unresolved／Koidzumi, Geniti 1939 Contributiones ad Cognitionem Florae Asiae Orientalis (continued from vol. VIII p. 61) Acta Phytotax. Geobot. 8 (2): 110
Rubus pungens auct. non Cambess.
Rubus rosaefolius Smith var. formosanus auct. non Cardot ／ Rubus rosaefolius Sm. [Spelling variant] → Rubus rosifolius Sm.
Rubus asper Wall. ex D. Don var. glaber (Koidz.) Hsieh → ???
Rubus sumatranus auct. non Miq.
Rubus pungens Cambess. var. oldhamii auct. non (Miq.) Maxim.
Rubus croceacanthus Lévl. var. glanduloso-calycinus (Hayata) S.-C. Liu → ???
:Rubus rosaefolius Smith subsp. maximowiczii Focke
果实近球形；花直径2-3厘米；小叶 (3)5-7枚，卵状椭圆形或卵状披针形，幼时两面密被柔毛，老时渐脱落，顶端急尖至短渐尖，边缘有不整齐锐锯齿或重锯齿；花常单生或2-5朵成伞房花序；萼片三角状披针形，长8-12毫米。
註:植物誌第一版：虎梅刺＝虎婆刺；毛刺懸鈎子　Rubus hirsutopungens Hayata

高砂懸鉤子　Rubus takasagoensis Koidz.

  • Rubus takasagoensis Koidz.；Koidzumi, Geniti 1939 Contributiones ad Cognitionem Florae Asiae Orientalis (continued from vol. VIII p. 61) Acta Phytotax. Geobot. 8 (2): 112
  • Rubus takasagoensis Koidz.；Ying, Col. Ill. Fl. Taiwan Intr. Pl., 台灣高等植物彩色圖誌 含引進種 2nd 481 1999



毛刺懸鈎子　Rubus hirsutopungens

Rubus hayatai Nemato ex Makino & Nemoto, Fl. Jap. ed. 2. 514. 1931.
Rubus hirsutus (Hayata) Hayata, l. c. 1: 227. 1911.
Rubus okamotoanus Koidz. in Acta Phytotax. Geobot. 8: 110. 1939.
Rubus rosaefolius Sm. var. hirsutus Hayata in Journ. Coll. Sci. Univ. Tokyo 25(19): 81. 1908 (Fl. Mont. Form.).

禿懸鉤子　Rubus croceacanthus var. glaber

梣葉懸鉤子　Rubus fraxinifoliolus

小叶常5枚，长圆披针形或披针形，两面无毛，下面沿中脉疏生小皮刺，边缘具不整齐单锯齿，花单生或成对，顶生或腋生；花梗长1.5-2厘米，无毛或近无毛；花萼外无毛或近无毛。

蘭嶼榿葉懸鉤子　Rubus fraxinifolius

Rubus fraxinifolius Poir. var. kotoensis (Hayata) Koidz. → Rubus fraxinifolius Poir.
Rubus kotoensis Hayata → Rubus fraxinifolius Poir.
Rubus alnifoliolatus Lévl. var. kotoensis (Hayata) Li → Rubus fraxinifolius Poir.
榿葉懸鉤子　Rubus alnifoliolatus Lévl. → Rubus alnifoliolatus H. Lévl. & Vaniot
小叶常7-9枚；花组成宽大圆锥花序；叶片卵状披针形至长椭圆状披针形，边缘具不规则锯齿或重锯齿；花梗长达1.2厘米；果实长圆形或椭圆形，长约1.2厘米。
  • Rubus fraxinifolius Poir.；Poir., Encyclopédie méthodique: Botanique 6: 242. 1806. (Encycl. Method. Bot.)；




  • Rubus fraxinifolius Poir. var. kotoensis (Hayata) Koidz., The Journal of the College of Science, Imperial University of Tokyo, Japan 34(2): 144. 1913. (J. Coll. Sci. Univ. Tokyo, )
  • Rubus alnifoliolatus Lévl. var. kotoensis (Hayata) Li, Woody Flora of Taiwan 320. 1963. (Woody Fl. Taiwan, )


  • Rubus alnifoliolatus Lévl.; Liu & Su, Flora of Taiwan 3: 107. 1977. (Fl. Taiwan, 台灣植物誌)




腺萼懸鉤子　Rubus glandulosocalycinus

臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11803
小叶3-7枚，卵状长圆形或卵状披针形，边缘有锐裂或缺刻状重锯齿；花常单生，稀2-5朵成伞房状花序；花直径2-5厘米；萼片长三角形或三角披针形。

蓬蘽　Rubus hirsutus

細葉懸鉤子　Rubus linearifoliolus

Rubus linearifoliolus Hayata ／ https://tai2.ntu.edu.tw/ebook/ebookpage.php?volume=7&book=Icon.%20Pl.%20Formos.&page=22
Rubus asper Wall. var. glaber (Koidz.) Hsieh
Rubus croceacanthus H. Lév. var. glaber auct. non Koidz.
Rubus tsangii Merr.
  • Rubus linearifoliolus Hayata；J.Y.Huang & J.M.Hu in Taiwania 54: 296 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310


  • Rubus rosaefolius Smith var. linearifolius (Hayata) Li, Woody Flora of Taiwan 315. 1963. (Woody Fl. Taiwan, )
  • Rubus asper Wall. var. glaber (Koidz.) Hsieh, Journal of Taiwan Museum 42: 106. 1989. pro parte.. (J. Taiwan Mus., 台灣省立博物館半年刊)
  • Rubus croceacanthus H. Lév. var. glaber auct. non Koidz.: Hsieh & Ohashi, Flora of Taiwan, second edition 3: 126. 1993. pro parte. (Fl. Taiwan 2nd edit., 台灣植物誌第二版)



Rubus tsangii Merr. var. linearifoliolus (Hayata) T.T.Yü et L.T.Lu；http://ylist.info/ylist_detail_display.php?pass=41948
Rubus tsangii Merr.；http://ylist.info/ylist_detail_display.php?pass=41947


  • Rubus tsangii Merr., Lingnan Sci. J. 13: 28 (1934).

小梣葉懸鉤子　Rubus × parvifraxinifolius

Rubus suzukianus Liu & Yang
Rubus suzukianus T. S. Liu & T. Y. Yang → 　梣葉懸鉤子　Rubus fraxinifoliolus Hayata
Rubus suzukianus Y. C. Liu & T. Y. Yang → 梣葉懸鉤子　Rubus fraxinifoliolus Hayata
Rubus leucanthus sensu Makino, Fl. Jap. Suppl. 350. 1936, non Hance.

刺莓　Rubus rosifolius

刺莓　Rubus taiwanianus Matsum. → Rubus rosifolius Sm.
腺斑懸鉤子　Rubus rosaefolius Smith ssp. maximowiczii sensu Hayata auct. non Focke → ???
腺斑懸鉤子　Rubus rosaefolius Sm. Var. tropicus sensu Makino → ???
腺斑懸鉤子　Rubus glandulosopunctatus Hayata → 刺莓　Rubus rosifolius Sm.
腺斑懸鉤子　Rubus parvirosaefolius Hayata ／ Rubus parvirosifolius Hayata → Unresolved
Rubus hopingensis Liu & Lu → Rubus rosifolius Sm.
Rubus lanyuensis Chang → 蘭嶼懸鉤子　Rubus lanyuensis Hung T. Chang
Rubus tagallus Cham. & Schl.
Rubus tagallus Cham. & Schltdl. → Rubus rosifolius Sm.
Rubus tagallus Cham. & Schl. var. lanyuensis (Chang) Ying → 蘭嶼懸鉤子　Rubus lanyuensis Hung T. Chang
薔薇葉懸鉤子　Rubus rosifolius Sm. var. formosanus Cardot → Rubus croceacanthus H. Lév. (China)
植株全体仅有柔毛，无腺毛；叶片宽卵形至椭圆状卵形，顶端圆钝，稀急尖，基部圆形或近心形；花梗长达2厘米；花直径达2.5厘米。
  • Rubus rosifolius var. linearifolius (Hayata) H.L.Li
Rubus linearifolius Hayata

蘭嶼懸鉤子　Rubus lanyuensis


紅腺懸鉤子　Rubus sumatranus

長果懸鉤子　Rubus dolichocephalus Hayata

絨毛葉組　Rubus Sect. Stimulantes

裡白懸鉤子　Rubus mesogaeus

Rubus rarissimus Hayata → Rubus mesogaeus Focke
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11810

紅梅消　Rubus parvifolius

Rubus triphyllus Thunb. → Rubus triphyllus Thunb., Syst. Veg., ed. 14 (J. A. Murray). 475 (1784). https://www.biodiversitylibrary.org/page/1479927#page/494/mode/1up
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11808
Rubus triphyllus Thunb. var. toapiensis Yamamoto ／Rubus triphyllus var. toapiensis Yamam. → Rubus parvifolius L. var. toapiensis (Yamam.) Hosok.
Rubus parvifolius L. var. concolor Makino & Nemoto subvar. subconcolor Masam. →
Rubus parvifolius L. var. toapiensis (Yamamoto) Hosok. → Rubus parvifolius L. var. toapiensis (Yamam.) Hosok.　台東紅梅消
Rubus parvifolius L. var. subconcolor (Cardot) Makino & Nemoto →

台東紅梅消　Rubus parvifolius var. toapiensis

圓錐序組　Rubus Sect. Thyrsidaei

小楤葉懸鉤子　Rubus parviaraliifolius

Rubus parviaraliifolius Hayata var. laxiflorus Liu & Lu ／ Rubus parviaraliifolius var. laxiflorus Y. C. Liu & F. Y. Lu → Rubus parviaraliifolius Hayata
臺灣植物圖譜 Icones Plantarum Formosanarumosa Vol. 7 https://tai2.ntu.edu.tw/ebook/ebookpage.php?book=Icon.%20Pl.%20Formos.&volume=7&list=11809

木莓组　Sec. Malachobatus　／單葉

錦葵葉組　Rubus Sect. Moluccani

羽萼懸鉤子　Rubus alceifolius

Rubus alceifolius Poiret → Rubus alceifolius Poir.

Rubus moluccanus auct. non L.
Rubus laciniato-stipulatus Hayata ／ Rubus laciniatostipulatus Hayata ex Koidz. → 新店懸鉤子　Rubus pinnatisepalus Hemsl.
新店懸鉤子　Rubus pinnatisepalus auct. non Hemsl.

灰葉懸鉤子　Rubus nagasawanus var. arachnoideus

Rubus nagasawanus Koidz. var. arachnoideus (Liu & Lu) Ying → ???
Rubus alceaefolia Poir. var. emigratus auct. non Focke
Rubus arachnoideus Liu & Lu → Rubus arachnoideus Y. C. Liu & F. Y. Lu
  • Rubus nagasawanus Koidz. var. arachnoideus (Liu & Lu) Ying；J.Y.Huang & J.M.Hu in Taiwania 54: 298 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310
  • Rubus arachnoideus Liu & Lu, Quart. J. Chin. For. 9: 126. fig. 1. 1976；Lu, Fu-Yuan 1976 Contributions to the dicotyledonous plants of Taiwan 2 Quart. J. Chin. For. 9(2): 127-143





Java and sumatra 首採

大同灰葉懸鉤子　Rubus nagasawanus

Rubus polyanthus Li
Rubus polyanthus H. L. Li → Rubus nagasawanus Koidz.
Rubus arachnoideus Y.-C. Liu & F.-Y. Lu var. tatongensis S.-C. Liu →

寒莓　Rubus buergeri

Rubus shimadai Hayata　／Rubus shimadae Hayata → Rubus buergeri Miq.
Rubus pseudo-buergeri Sasaki　／假寒莓　Rubus pseudobuergeri Sasaki → Rubus buergeri Miq.
Rubus buergeri Miq. var. pseudo-buergeri (Sasaki) Liu & Yang →

托葉和苞片掌狀或羽狀較淺裂。 矮小攀援或匍匐灌木；葉背絨毛老時脫落；花數朵成頂生短總狀花序或數朵簇生葉腋或單生。 葉背密被絨毛。 枝、葉柄和花序被絨毛狀長柔毛；葉邊裂片圓鈍；花萼密被淡黃色長柔毛和絨毛；萼片披針形或卵狀披針形，外萼片僅頂端淺裂。

台灣懸鉤子　Rubus formosensis

Rubus formosanus Maxim. ex Focke → Rubus formosensis Kuntze
Rubus randaiensis Hayata → Rubus formosensis Kuntze
Rubus nantoensis Hayata → Rubus formosensis Kuntze
Rubus rugosissimus Hayata → Rubus formosensis Kuntze

湖南懸鉤子　Rubus hunanensis

葉近圓形或寬 卵形，頂端急尖，基部深心形，枝、葉柄和花序被細短柔毛；葉邊裂片急尖；花萼被灰白色至黃灰色短柔毛和絨毛；萼片寬卵形，外萼片邊緣羽狀條裂。


新店懸鉤子　Rubus pinnatisepalus

清水懸鉤子　Rubus pseudoacer subsp. flexuosus

Rubus pseudoacer Makino var. flexuosus Liu & Lu → Rubus trifidus Thunb.
  • Rubus pseudoacer Makino var. flexuosus Liu & Lu, Quart. J. Chin. For. 9(2): 133. fig. 5. 1976；Lu, Fu-Yuan 1976 Contributions to the dicotyledonous plants of Taiwan 2 Quart. J. Chin. For. 9(2): 127-143


棕紅懸鉤子　Rubus rufus

Rubus yuliensis Liu & Lu → Rubus yuliensis Y. C. Liu & F. Y. Lu　玉里懸鉤子
Rubus rufus Focke var. yuliensis (Y.-C. Liu & F.-Y. Lu) S.-C. Liu →
  • Rubus rufus Focke；J.Y.Huang & J.M.Hu in Taiwania 54: 304 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310


  • Rubus yuliensis Y.C.Liu & F.Y.Lu；Lu, Quart. J. Chin. Forest. 9(2): 135 1976；Lu, Fu-Yuan 1976 Contributions to the dicotyledonous plants of Taiwan 2 Quart. J. Chin. For. 9(2): 127-143
  • Rubus rufus Focke var. yuliensis (Y.-C. Liu & F.-Y. Lu) S.-C. Liu, 1: 130. 2000. (Trees Taiwan, )



玉里懸鉤子　Rubus yuliensis



尖葉組　Rubus Sect. Acuminati

单叶，宽卵形至长圆形，顶端渐尖至尾尖，有网状脉或基部具5出脉，叶片下面无毛或有柔毛，边缘不分裂或浅裂；托叶和苞片掌状或羽状条裂，或不分裂；花组成顶生宽大或狭圆锥花序，腋生花序较小，有时近总状；心皮约10-40

高梁泡　Rubus lambertianus

Rubus lambertianus Ser. ssp. xanthoneurus auct. non Focke

尾葉懸鉤子　Rubus lambertianus var. morii

Rubus lambertianus Ser. var. morii (Hayata) S. S. Ying →

Rubus morii Hayata → 尾葉懸鉤子　Rubus lambertianus Ser. var. glandulosus Cardot
Rubus gelatinosus Sasaki → unresolved
Rubus tiponensis Hosokawa → unresolved
Rubus morii Hayata var. urophyllus (Y.-C. Liu & F.-Y. Lu) S.-C. Liu →
Rubus urophyllus Liu & Lu → 長尾葉懸鉤子　Rubus urophyllus Y. C. Liu & F. Y. Lu → unresolved

長尾葉懸鉤子　Rubus urophyllus

Rubus hakonensis

Rubus chaetophorus

長葉組　Rubus Sect. Dolichophylli

裂葉懸鉤子　Rubus howii

  • Rubus howii Merr. & Chun, Sunyatsenia 5: 71 T. 8 (1940).


  • Rubus howii Merr. & Chun；J.Y.Huang & J.M.Hu in Taiwania 54: 294 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310



  • Chen, Po-Hao, Juinn-Yih Huang and An-Ching Chung 2024 Rubus puyumaensis, A New Species of Rosaceae from Taiwan Biodiversity Data Journal 12: e115431.
  • Rubus howii auct. non Merr. & Chun

長序組　Rubus Sect. Elongati

单叶，卵形、长卵形、卵状披针形至长圆形，稀宽卵形或近圆形，顶端渐尖，稀急尖或圆钝，叶片下面密被灰白色至黄灰色绒毛，基部常有5出脉，边缘不分裂或浅裂；托叶和苞片不分裂、浅裂或深裂；花组成顶生宽大圆锥花序，花序枝长大而开展呈金字塔形，长可达27厘米，稀为狭窄圆锥花序，腋生花序较小；心皮约10-50。

粗毛懸鉤子　Rubus tephrodes var. setosissimus

Rubus nagasawanus auct. non Koidz.
Rubus tephrodes auct. non Hance
Rubus alceaefolia Poir. Var. emigratus sensu Koidz. → ????
Rubus formosensis sensu Matsum. → 粗毛懸鉤子　Rubus nagasawanus Koidz.／ In Bot. Mag. Tokyo 15: 156. 1901, non O. Ktze.
Rubus polyanthus Li → 粗毛懸鉤子　Rubus nagasawanus Koidz.
Rubus tephrodes Hance var. setosissinus sensu Koidz. In Acta Phytotax. Geobot. 8: 262. 1939, non Hand-Mazz.

灰白毛莓　Rubus tephrodes


鏽葉組　Rubus Sect. Fuscifolii

裂緣苞懸鉤子　Rubus flagelliflorus


  • Rubus flagelliflorus FockeJ.Y.Huang & J.M.Hu in Taiwania 54: 292 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310


胡氏懸鉤子 Rubus reflexus var. hui

Rubus hui Diels → Rubus reflexus Ker Gawl. var. hui (Diels ex Hu) F. P. Metcalf

銹毛莓　Rubus reflexus

梨葉組　Rubus Sect. Pirifolii

葉片卵形、長圓形至長圓披針形，頂端急尖或短漸尖；葉柄長約1釐米，極稀達2釐米；心皮5-10。

梨葉懸鉤子　Rubus pyrifolius

Rubus floribundo-paniculatus Hayata ／ Rubus floribundopaniculatus Hayata → Rubus pirifolius Sm.
Rubus parvipetalus Odashima → Rubus pirifolius Sm.

總序組　Rubus Sect. Sozostyli

桑葉懸鉤子　Rubus kawakamii

桑葉懸鉤子　Rubus swinhoei Hance var. kawakamii (Hayata) S.-C. Liu → Rubus kawakamii Hayata

斯氏懸鉤子　Rubus swinhoei

Rubus adenotrichopodus Hayata → Rubus swinhoei Hance

柳氏懸鉤子　Rubus liui

Rubus ilanensis Liu & Lu → Rubus liui Yuen P. Yang & S. Y. Lu

雜交種

  • Rubus croceacanthus × corchorifolius
  • Rubus rosifolius × fraxinifolius
  • J.Y.Huang & J.M.Hu in Taiwania 54: 308 (2009)／Huang, Juinn-Yih and Jer-Ming Hu 2009 Revision of Rubus (Rosaceae) in Taiwan Taiwania 54(4): 285-310

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台、日、中

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Prunus／Cerasus Reference

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