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Rubiaceae Reference


  1. Alejandro, G. D. and S. Liede 2003 The Philippine Rubiaceae genera: updated synopsis in intkey databases of the delta system Blumea 48 (2): 261-277
  2. Alejandro, Grecebio Jonathan D. 2007 The Current Status of the Philippine Rubiaceae Philippine Journal of Systematic Biology 1 (1)
  3. Andreasen, Katarina and Birgitta Bremer 2000 Combined phylogenetic analysis in the Rubiaceae-Ixoroideae: morphology, nuclear and chloroplast DNA data American Journal of Botany 87(11):1731-1748
  4. Arriola, A. H., P. D. Camacho, M. J. A. Calaramo and G. J. D. Alejandro 2015 Pyrostria triflora , a new species of Vanguerieae (Rubiaceae) from Luzon, Philippines Bangladesh J. Plant Taxon. 22 (1): 55–58
  5. Bakhuizen van den Brink, Jr., R. C. 1975 A Synoptical Key to the Genera of the Rubiaceae of Thailand Thai Forest Bulletin (Botany) 9: 15-55
  6. Banag, Cecilia Illiscupides, Danilo Tandang, Ulrich Meve, Sigrid Liede-Schumann 2015 Two new species of Ixora (Ixoroideae, Rubiaceae) endemic to the Philippines Phytotaxa 202 (2): 155–160
  7. Banag, Cecilia I., Arnaud Mouly, Grecebio Jonathan D. Alejandro, Birgitta Bremer, Ulrich Meve, Guido W. Grimm and Sigrid Liede-Schumann 2017 Ixora (Rubiaceae) on the Philippines - crossroad or cradle? BMC Evolutionary Biology 17:131
  8. Bremekamp, C. E. B. 1940 On Urophyllum Wall. (Rubiaceae) and its nearest allies Recueil des Travaux Botaniques Neerlandais 171-197
  9. Bremer, Birgitta 1989 The genus Argostemma (Rubiaceae-Argostemmateae) in Borneo Annals of the Missouri Botanical Garden 76 (1) 7-49
  10. Bremer, Birgitta 2009 A Review of Molecular Phylogenetic Studies of Rubiaceae 1 of the Missouri Botanical Garden 96 (1):4-26
  11. Bremer, Birgitta and Torsten Eriksson 2009 Time tree of Rubiaceae: phylogeny and dating the family, subfamilies, amd tribes Int. J. Plant Sci. 170(6):766–793
  12. Chang, Ching-en 1981 An Eumeration of the Woody Plants of Botel Tobago J. Phytogeogr. Taxon. 29 (1): 1-21
  13. Chang, Kun-Cheng, Yen-Hsueh Tseng, Chin-Chiang Wang, Shu-Lin Deng, Chiu-Mei WANG 2008 Note on Neanotis (Rubiaceae) in Taiwan. Taiwan Journal of Forestry Science 23(3): 271-277.
  14. Chantaranothai, Pranom 2018 Nomenclatural notes on the genus Schizomussaenda H. L. Li (Rubiaceae) Thai For. Bull (Bot.) 46 (1): 58-61
  15. Chao, Jew-Ming 1966 The Taiwan species of Rubia (Rubiaceae) BioFormosa 1:46-51
  16. Chavez1, Jayson G., Ulrich Meve and Grecebio Jonathan D. Alejandro1 2012 New Combinations in Rothmannia (Rubiaceae) and the Establishment of the Genus in the Philippine Flora Philippine Science Letters 5 (2): 216-223
  17. Chaw, S. M., V. V. Sivarajan 1989 Seed coat micromorphology of some Asiatic Spermacoceae (Rubiaceae) Bot. Bull. Academia Sinica 30(1) 15-24
  18. Chen, Junhao, Khoon Meng Wong, Antony van der Ent and Hugh Tiang Wah Tan 2014 Nine new species of Timonius (Rubiaceae) from Kinabalu Park, Borneo Phytotaxa 181 (3): 138–150
  19. Chen, Junhao, Hugh T. W. Tan and K. M. Wong 2015 A revision of Timonius (Rubiaceae) in Kinabalu Park, Borneo, with notes on typification and species distinction Plant Ecology and Evolution 148 (3): 420–430
  20. Chou, Fu-Shan, Chih-Kai Yang and Chun-Kuei Liao 2006 Taxonomic Status of Ophiorrhiza michelloides (Masam.) X. R. Lo (Rubiaceae) in Taiwan Taiwania 51(2): 143-147
  21. Church, Sheri A. and Douglas R. Taylor. 2005 Speciation and hybridization among Houstonia (Rubiaceae) species: The influence of polyploidy on reticulate evolution American Journal of Botany 92(8): 1372–1380
  22. Corrêa, Andréa Macêdo and Eliana Regina Forni-Martins 2004 Chromosomal studies of species of Rubiaceae (A. L. de Jussieu) from the Brazilian cerrado CARYOLOGIA 57(3) 250-258
  23. Davis, Aaron P., Diane Bridson, Charlie E. Jarvis and Rafaël Herman Anna Govaerts 2001 The typification and characterization of the genus Psychotria L. (Rubiaceae) Botanical Journal of the Linnean Society 135 (1): 35–42
  24. Davis, Arron P., Rafel Govaerts, Diane M. Bridson and Piet Sstoffelen 2006 An annotated taxonomic conspectus of the genus Coffea (Rubiaceae) Botanical Journal of the Linnean Society 152 465–512
  25. Delprete, Piero G. and Joseph H. Kirkbride, Jr. 2008 Clarification of Borreria gymnocephala, Diodia gymnocephala, Diodia schumannii, Borreria flavovirens, and Spermacoce schumannii (Rubiaceae) Journal of the Botanical Research Institute of Texas 2(1) 305-308
  26. Dessein, Steven, Helga Ochoterena, Petra De Block, Frederic Lens, Elmar Robbrecht, Peter Schols, Erik Smets, Stefan Vinckier, and Suzy Huysmans 2005 Palynological Characters and Their Phylogenetic Signal in Rubiaceae The Botanical Review 71(3):354-414
  27. Ehrendorfer, Friedrich 2010 New and Critical Taxa of Rubia and Galium (Rubiaceae, Rubieae) for the Flora of China Novon 20(3):268-277
  28. Gao, Yi-Ding, Rui-Jiang Wang and Ching-I Peng 2010 Diodia teres Walt. (Rubiaceae), a Newly Recorded Weed in Fujian Taiwania 55(2): 177-179
  29. Groeninckx, Inge, Steven Dessein, Helga Ochoterena, Claes Persson, Timothy J. Motley, Jesper Kårehed, Birgitta Bremer, Suzy Huysmans, and Erik Smets 2009 Phylogeny of the Herbaceous Tribe Spermacoceae (Rubiaceae) Based on Plastid DNA Data Annals of the Missouri Botanical Garden 96(1):109-132
  30. Heng, Li 1999 A New Species of Paederia (Rubiaceae) from China Novon 9(2): 220
  31. Holm, Theo. 1907 Rubiaceae: Anatomical Studies of North American Representatives of Cephalanthus, Oldenlandia, Houstonia, Mitchella, Diodia, and Galium Botanical Gazette 43(3) 153-186
  32. Hsieh, Chang-Fu and Shu-Miaw Chaw 1987 Diodia virginiana L. (Rubiaceae) in Hsinchu: New to Taiwan Bot. Bull. Academia Sinica 28(1) 43-48
  33. Hsieh, Tsung-Hsin and Po-Yi Chiang 2010 Galium paradoxum Maxim. (Rubiaceae), a newly recorded plant in Taiwan Taiwania 55(2) 192-194
  34. Hsu, Chien-Chang 1973 Some Noteworthy Plants Found in Taiwan Taiwania 18(1) 62-72 / Gomphrena celosioidesCoronopus didymusCuphea carthagenensisPlumbago zeylanicaBorreria latifoliaBorreria laevisAmbrosia elatiorGaillardia pulchellaGalinsoga parvifloraSenecio vulgarisSolidago altissimaNajas marinaNajas minorNajas indicaPanicum dichotomiflorumPaspalum virgatum
  35. Hsu, Tian-Chuan, and Zhi-Hao Chen 2017 Scleromitrion sirayanum (Rubiaceae: Spermacoceae), a new species of the Hedyotis-Oldenlandia complex in Taiwan Taiwania 62(2): 151‒156
  36. Huang S. F. and T.C. Huang. 1997 Notes on the Flora of Taiwan (27) The Genus Argostemma Wall.(Rubiaceae). Taiwania 42(1): 1-7.
  37. Huang, Tseng-Chieng, Shing-Fan Huang, Kuoh-Cheng Yanf 1989 Notes on the Flora of Taiwan (3)--- The miscellaneous plants Taiwania 34(1) 45-53 / Epithema brunonis var. fasciculataTheligonum formosanumRotala wallichiiEdsmanthus virgatusandParis fargesii var. brevipetalata
  38. Huxley, Camilla R. 1978 The Ant-Plants Myrmecodia and Hydnophytum (Rubiaceae), and the relationship between their morphology, ant occupants, physiology and ecology New Phytol. 80 231-268
  39. Johansson, J. T. 1989 Revision of the genus Rennellia Korth. (Rubiaceae-Rubioideae) Blumea 34 3-19
  40. Judkevich, Marina D., Roberto M. Salas and Ana M. Gonzalez 2017 Colleters in American Spermacoceae Genera (Rubiaceae): Morphoanatomical and Evolutionary Aspects Int. J. Plant Sci. 178 (5): 378–397
  41. Jung, Ming-Jer, Shin-Ming Ku and Ching-I Peng 2011 Oldenlandiopsis Terell & W. H. Lewis (Rubiaceae), a Newly Recorded Genus in Taiwan Taiwania 56(1) 58-61
  42. Kårehed, Jesper and Birgitta Bremer 2007 The systematics of Knoxieae (Rubiaceae)—molecular data and their taxonomic consequences Taxon 56 (4): 1051-1076
  43. Kåreheda,, Jesper, Inge Groeninckxb, Steven Desseinc, Timothy J. Motleyd, Birgitta Bremer 2008 The phylogenetic utility of chloroplast and nuclear DNA markers and the phylogeny of the Rubiaceae tribe Spermacoceae Molecular Phylogenetics and Evolution 49(3) 843–866
  44. Kesonbua, Wirot and Pranom Chantaranothai 2008 A checklist of the genus Tarenna Gaertn. (Rubiaceae) in Thailand THAI FOR. BULL. (BOT.) 36: 18–45
  45. Kesonbua, Wirot and Pranom Chantaranothai 2013 The genus Morinda (Rubiaceae) in Thailand ScienceAsia 39: 331–339
  46. Khushbu, Chauhan, Patel Anar, Patel Mayuree, Macwan Carol, Solanki Roshni, Adeshara Subodh. 2010 Paederia foetida Linn. As a potential medicinal plant : A review Journal of Pharmacy Research 3(12): 3135-3137
  47. Kitagawa, Masao 1934 Contributio ad Cognitionem Florae Manshuricae IV Bot. Mag. (Tokyo) 48 (573): 610-620
  48. Kliphuis, E. 1970 Cytotaxonomic notes on some Galium species, Galium silvaticum L., Galium aristatum L. and Galium schultesii Vest Botanical Museum and Herbarium 271-283
  49. Kliphuis E. 1983 Cytotaxonomic notes on some species of the gneus Galium collected in the Spain Lagascalia 11(2): 229-244
  50. Kliphuis E. 1984 Cytotaxonomie studies on Galium verum L. Botanica Helvetica 94/2 269-278
  51. Koek-Noorman, Jifke 1972 The wood anatomy of Gardenieae, Ixoreae and Mussaendeae (Rubiaceae) Acta Bot. Neerl. 21 (3): 301-320
  52. Kuanga, Yan-Feng, Bruce K. Kirchoff, Yuan-Jiang Tang, Yuan-Hui Liang, and Jing-Ping Liao. 2008. Palynological characters and their systematic significance in Naucleeae (Cinchonoideae, Rubiaceae). Review of Palaeobotany and Palynology 151(3-4): 123-135
  53. Lewis, Walter H. and Royce L. Oliver 1974 Revision of Richardia (Rubiaceae) Brittonia 26(3):271-301
  54. Lin, Chang-Jung and Chien-Fan Chen 2013 Mitracarpus hirtus (L.) DC. (Rubiaceae), a Newly Naturalized Plant in Taiwan TW J. of Biodivers. 15 (2): 155 -161
  55. Low, Y. W. and K. M. Wong 2007 Two new species of Gardenia (Rubiaceae) from Borneo and notes on Gardenia pterocalyx Edinburgh Journal of Botany 64 (1): 25-36
  56. Low, Yee Wen, John B. Sugau and Khoon Meng Wong 2016 Hydnophytum puffii (Rubiaceae: Psychotrieae), a new ant-plant from Sabah, Malaysian Borneo Plant Ecology and Evolution 149 (1): 123–130
  57. Makino, T. 1903 Observations on the Flora of Japan. (Continued from p. 62.) Bot. Mag. Tokyo 17 (194): 70-80
  58. Malcomber, Simon T. 2002 Phylogeny of Gaertnera Lam. (Rubiaceae) based on multiple DNA markers: evidence of a rapid radiation in a widespread, morphologically diverse genus [Evolution] 56 (1): 42–57
  59. Manns, Ulrika, Niklas Wikström, Charlotte M. Taylor and Birgitta Bremer 2012 Historical biogeography of the predominantly neotropical subfamily Cinchonoideae (Rubiaceae): into or out of America? Int. J. Plant Sci. 173(3):261–289
  60. Mikhama, K. and P. Sirisant 2016 The last and largest of Cephalanthus tetrandra freshwater swamp forest in Northeast Thailand: Natural resource appreciation and management of local community International Journal of Agricultural Technology 12 (3): 429-438
  61. Mouly, Arnaud, Sylvain G. Razafimandimbison, Anbar Khodabandeh and Birgitta Bremer 2009 Phylogeny and classification of the species-rich pantropical showy genus Ixora (Rubiaceae-Ixoreae) with indications of geographical monophyletic units and hybrids American Journal of Botany 96(3): 686–706


  1. Nakamura, Koh, Shih-wen Chung, Goro Kokubugata, Tetsuo Denda, Masatsugu Yokota 2006 Phylogenetic systematics of the monotypic genus Hayataella (Rubiaceae) endemic to Taiwan J. Plant Research 119: 657-661
  2. Ordas, Jorge Anton D., Neil Alejandro A. Pinarok, Roanne B. Romeroso, Grecebio Jonathan D. Alejandro, Cecilia I. Banag-Moran 2019 A checklist of Rubiaceae species from Eastern Samar, Visayas, Philippines Check List 15 (2): 295–312
  3. Ou, Chern-hsiung 1987 Contributtons to the dicotyledonous plants of Taiwan (X) Bullitin Exper. for NCHU 8 11-30
  4. Puff, C., J. Greimler and R. Buchner 1998 Revision of Schradera (Rubiaceae-Schradereae) in Malesia Blumea 43 287-335
  5. Puff, Christian 2001 The Significance of Gynoecium and Fruit and Seed Characters for the Classification of the Rubiaceae Malayan Nature Joumal 55 (1 & 2): 133-146
  6. Puff, christian 2010 Mycetia (Rubiaceae) from Khao Soi Dao, Chanthaburi Province, SE Thailand: the discovery of a remarkable new species and the true identity of a supposedly endemic species Thai. For. Bull. (Bot.) 38: 173–178
  7. Purwantoto, R. Sibekti, Harry Wiriadinata and Susiani Purbaningsih 2005 Gatra Taksonomi Argostemma Wall. (Rubiaceae-Rubioidae) di Gunung Gede-Pangrango, Gunung Halimun, dan Gunung Salak berdasarkan Karakter Morfologi Biodiveritas 6 (3) 199-204
  8. Rajaseger, G., H. T. W. Tan, I. M. Turner and P. P. Kumar 1997 Analysis of Genetic Diversity among Ixora Cultivars (Rubiaceae) using Random Amplified Polymorphic DNA Annals of Botany 80: 355–361
  9. Raju, Aluri Jacob Solomon and Janga Radhakrishna 2017 Contribution to the knowledge of three Indian Spermacoce L. (Rubiaceae) and some preliminary information about their pollination ecology Anales de Biología 39: 111-126
  10. Razafimandimbison, Sylvain G. and Birgitta Bremer 2002 Phylogeny and classification of Naucleeae s.l. (Rubiaceae) inferred from molecular (ITS, rBCL, and tRNT-F) and morphological data American Journal of Botany 89(7): 1027-1041
  11. RazaWmandimbison, Sylvain G., Joachim Moog, Henrik Lantz, Ulrich Maschwitz, Birgitta Bremera 2005 Re-assessment of monophyly, evolution of myrmecophytism, and rapid radiation in Neonauclea s.s. (Rubiaceae) Molecular Phylogenetics and Evolution 34(2): 334–354
  12. Razafimandimbisona, Sylvain G., Timothy D. McDowell, David A. Halfordc, Birgitta Bremerb 2009 Molecular phylogenetics and generic assessment in the tribe Morindeae (Rubiaceae–Rubioideae): How to circumscribe Morinda L. to be monophyletic? Molecular Phylogenetics and Evolution 52(3) 879-886
  13. Razafimandimbison, Sylvain G, Stefan Ekman, Timothy D. McDowell and Birgitta Bremera 2012 Evolution of growth habit, inflorescence architecture, flower size, and fruit type in Rubiaceae: Its ecological and evolutionary implications PLoS ONE 7(7): e40851
  14. Ridsdale, C. E. 1976 A revision of the tribe Cephalantheae (Rubiaceae) Blumea 23 (1): 177–188.
  15. Ridsdale, C.E. (1978). A revision of Mitragyna and Uncaria (Rubiaceae) Blumea 24(1), 43–100
  16. Ridsdale, C. E. 1978 A revision of the Tribe Naucleeae s.s. (Rubiaceae) Blumea 24 307-356
  17. Ridsdale, C. E. 1989 A revision of Neonauclea (Rubiaceae) Blumea 34 (1): 177–275
  18. Ridsdale, C. E. 2008. Thorny problems in the Rubiaceae: Benkara, Fagerlindia and Oxyceros Reinwardtia 12(4):289–298
  19. Robbrecht, Elmar 1988 Tropical woody Rubiaceae Volume 1 National Botanic Garden of Belgium ISBN 90-72619-02-1
  20. Rydin, Catarina, Sylvain G. Razafimandimbison, Anbar Khodabandeh and Birgitta Bremer 2009 Evolutionary relationships in the Spermacoceae alliance (Rubiaceae) using information from six molecular loci: insights into systematic affinities of Neohymenopogon and Mouretia. Taxon 58(3):793-810
  21. Schanzer, Ivan A. 2004 Systematic notes on Ophiorrhiza trichocarpon Blume (Rubiaceae) and some related species THAI FOR. BULL. (BOT.) 32: 132–145
  22. Seah, W. W. and K. M. Wong 2018 Flora of Singapore precursors, 1. Gynochthodes praetermissa (Rubiaceae: Morindeae), a new West Malesian species, with notes on related taxa Gardens' Bulletin Singapore 70 (2): 267–273
  23. Sivadasan, M. and N. Mohanan 1991 Ixora agasthyamalayana, a new species of Rubiaceae from India Bot. Bull. Acad. Sin. 32(4): 323-316
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  25. Soza, Valerie L. & Richard G. Olmstead 2010 Molecular systematics of tribe Rubieae (Rubiaceae): Evolution of major clades, development of leaf-like whorls, and biogeography TAXON 59(3) 2010: 755–771
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  30. Terrell, Edward E. 1990 Synopsis of Oldenlandia (Rubiaceae) in the United States Phytologia 68(2):125-133
  31. Terrell, Edward E. and Harold Robinson 2009 A new genus, Mexotis, for five mexican species of Hedyotideae (Rubiaceae) J. Bot. Res. Inst. Texas 3(1): 59–70
  32. Tosh, James, Aaron P. Davis, Steven Dessein, Petra De Block, Suzy Huysmans, Mike F. Fay, Erik Smets, and Elmar Robbrecht 2009 Phylogeny of Tricalysia (Rubiaceae) and its Relationships with Allied Genera Based on Plastid DNA Data: Resurrection of the Genus Empogona Annals of the Missouri Botanical Garden 96(1):194-213
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  1. 段林东、林祁 2007 中国蛇根草属(茜草科)几种植物的分类学订正 植物分类学报 45(6):870-879
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  3. 陈伟球 1983 中国水锦树属的初步研究 植物分类学报 21(3): 277-284
  4. 陈伟球 1983 中国水锦树属的初步研究(续) 植物分类学报 21(4): 386-403
  5. 御江久夫 1938 雑録--びらうどへくそかづらノ學名 植物研究雑誌 14 (4):280
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HedyotisOldenlandia Reference

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  2. Chen, Shih-Huei, Jian-Yu Su and Ming-Jou Wu 2010 Hedyotis pinifolia Wall. ex G. Don (Rubiaceae), a New Record to the Flora of Taiwan Taiwania 55(1): 86-89, 2010
  3. Florentin, Mariela Nuñez, Andrea Cabaña Fader and Ana Maria Gonzalez 2016 Morpho-anatomical and morphometric studies of the floral structures of the distylous Oldenlandia salzmannii (Rubiaceae) Acta Botanica Brasilica
  4. Fukuoka, Nobuyuki 1970 Contributions to the Flora of Southeast Asia III : Hedyotis (Rubiaceae) of Thailand Tonan Ajia Kenkyu 8(3): 305-336
  5. Guo, Xing, Mark P. Simmons, Paul Pui-Hay But, Pang-Chui Shaw, Rui-Jiang Wang 2011 Application of DNA barcodes in Hedyotis L. (Spermacoceae, Rubiaceae) Journal of Systematics and Evolution 49(3) 203-212
  6. Guo, Xing, Rui-Jiang Wang, Mark P. Simmons, Paul Pui-Hay But, Jing Yu 2013 Phylogeny of the Asian HedyotisOldenlandia complex (Spermacoceae, Rubiaceae): Evidence for high levels of polyphyly and the parallel evolution of diplophragmous capsules Molecular Phylogenetics and Evolution 67 110–122
  7. Hsu, Tian-Chuan, Zhi-Hao Chen, Shih-Wen Chung 2014 Hedyotis koana Wang (Rubiceae) a Newly Recorded Plant in Taiwan Quarterly Journal of Forest Research 36(3):179-182
  8. Hsu, Tian-Chuan and Zhi-Hao Chen 2017 Scleromitrion sirayanum (Rubiaceae: Spermacoceae), a new species of the Hedyotis-Oldenlandia complex in Taiwan Taiwania 62(2): 151‒156
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  12. Sivarajan, V. V., S. D. Biju and Philip Mathew 1993 Taxonomy and Nomenclature of Some Indian Species of Hedyotis (Rubiaceae) Kew Bulletin 48 (2):391
  13. Terrell, Edward E. and Harold Robinson 2010 Taxonomic notes on the genera Stenotis and Carterella (Rubiaceae) and transfer of Hedyotis greenei to Stenotis J. Bot. Res. Inst. Texas 4(2):619–623.
  14. Wang, Gang-Tao, Ying Zhang, Dan Liang, Yu-Qiang Chen and Rui-Jiang Wang 2018 Hedyotis taishanensis (Rubiaceae): A new species from Guangdong, China Phytotaxa 367 (1): 38-44
  15. Wang, Rui-Jiang 2007 Hedyotis koana R. J. Wang, a new species of Rubiaceae from China Acta Phytotaxonomica Sinica 45(5): 696-700
  16. Wang, Rui-Jiang 2008 Two New Species of Hedyotis (Rubiaceae: Hedyotideae) from Hainan, China Novon 18: 264-268
  17. Wang, Rui-Jiang, Shu-jun Deng, Qi Liao 2014 Nomenclature Clarification of the Traditional Chinese Medicine Baihuasheshecao and Its Adulterants Based on Molecular and Morphological Evidence J. Trop. Subtrop. Bot. 22 (5): 431-442
  18. Wikström, Niklas, Suman Neupane, Jesper Kårehed, Timothy J. Motley and Birgitta Bremer 2013 Phylogeny of Hedyotis L. (Rubiaceae: Spermacoceae): Redefining a complex Asian-Pacific assemblage TAXON 62 (2): 357–374

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  2. 陈雨晴,郭兴,王瑞江 2015 中国耳草属(茜草科)植物新记录种 热带亚热带植物学报 23 (1): 31 ~ 33

Lasianthus Reference

  1. Cai, Min, Hua Zhu, Hong Wang 2008 Pollen morphology of the genus Lasianthus (Rubiaceae) and related taxa from Asia Journal of Systematics and Evolution 46(1): 62–72
  2. Liu, Tang-Shui and Jew-Ming Chao 1964 Revision of the Taiwan species of Lasianthus (Rubiaceae) Taiwania 10:117-150
  3. Matsumura, J. 1901 Notulæ ad Plantas Asiaticas Orientales (Continued from p. 147.) Bot. Mag. Tokyo 15 (168): 13-17
  4. Matsumura, J. 1901 Notulæ ad Plantas Asiaticas Orientales (Continued from p. 117.) Bot. Mag. Tokyo 15 (169): 37-41
  5. Xiao, Long-Qian and Hua Zhu 2007 Paraphyly and phylogenetic relationships in Lasianthus (Rubiaceae) inferred from chloroplast rps16 data Bot. Stud. 48:227-232
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  7. Zhu, Hua 2001 A taxonomic revision of the genus Lasianthus Jack. (Rubiaceae) from Thailand Acta Phytotaxonomica Sinica 39(2):116-150
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  9. Chiou, Chyi-Rong, Guo-Zhang Michael Song, Jui-Han Chien, Chang-Fu Hsieh, Jenn Che Wang, Ming-Yih Chen, Ho-Yih Liu, Ching-Long Yeh, Yue-Joe Hsia, and Tze-Ying Chen 2010 Altitudinal distribution patterns of plant species in Taiwan are mainly determined by the northeast monsoon rather than the heat retention mechanism of Massenerhebung Botanical Studies 51: 89-97
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  19. Editorial Commitee fo the Flora of Taiwan 2000 Flora of Taiwan, Second edition. Volume Five, Taiwan University, Taipei, Taiwan 106, the Republic of China, ISBN 957-9019-92-4
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